The Gastropoda or gastropods, more commonly known as snails and slugs, are a large taxonomic class within the Mollusca. The class Gastropoda includes snails and slugs of all kinds and all sizes from microscopic to quite large. There are huge numbers of sea snails and sea slugs, as well as freshwater snails and freshwater limpets, and land snails and land slugs.
The class Gastropoda contains a vast total of named species, second only to the insects in overall number. The fossil history of this class goes all the way back to the Late Cambrian. There are 611 families of gastropods, of which 202 families are extinct, being found only in the fossil record.
Gastropoda (previously known as univalves and sometimes spelled Gasteropoda) are a major part of the phylum Mollusca and are the most highly diversified class in the phylum, with 60,000 to 80,000 living snail and slug species. The anatomy, behavior, feeding and reproductive adaptations of gastropods vary significantly from one clade or group to another. Therefore, it is difficult to state many generalities for all gastropods.
The class Gastropoda has an extraordinary diversification of habitats. Representatives live in gardens, in woodland, in deserts, and on mountains; in small ditches, great rivers and lakes; in estuaries, mudflats, the rocky intertidal, the sandy subtidal, in the abyssal depths of the oceans including the hydrothermal vents, and numerous other ecological niches, including parasitic ones.
Although the name "snail" can be, and often is, applied to all the members of this class, commonly this word means only those species with an external shell large enough that the soft parts can withdraw completely into it. Those gastropods without a shell, and those with only a very reduced or internal shell, are usually known as slugs.
The marine shelled species of gastropod include edible species such as abalone, conches, periwinkles, whelks, and numerous other sea snails that produce seashells which are coiled in the adult stage, even though in some cases the coiling may not be very visible, for example in cowries. There are also a number of families of species such as all the various limpets, where the shell is coiled only in the larval stage, and is a simple conical structure after that.
The word "gastropod" is derived from the Ancient Greek words ''γαστήρ (gastér, ''stem:'' gastr-)'' "stomach", and ''πούς (poús, ''stem:'' pod-)'' "foot", hence stomach-foot. This is an anthropomorphic name, based on the fact that to humans it appears as if snails and slugs crawl on their bellies. In reality, snails and slugs have their stomach, the rest of their digestive system and all the rest of their viscera in a hump on the opposite, dorsal side of the body. In most gastropods this visceral hump is covered by, and contained within, the shell.
In the scientific literature, gastropods were described under the vernacular (French) name "gasteropodes" by Georges Cuvier in 1795. The name was later Latinized.
The earlier name ''univalve'' means "one valve" or shell, in contrast to ''bivalve'' applied to mollusks such as clams and meaning that those animals possess two valves or shells.
Gastropods are the class of molluscs which have the greatest numbers of named species. However estimates of the total number of gastropod species varies widely, depending on the cited sources. The number of gastropod species can be deduced from estimates of the number of described species of Mollusca with accepted names: about 85,000 (minimum 50,000, maximum 120,000). But an estimate of the total number of Mollusca, including undescribed species, is about 240,000 species. The estimate of 85,000 molluscs includes 24,000 described species of terrestrial gastropods.
Different estimations (from different sources) for aquatic gastropods give about 30,000 species of marine gastropods and about 5,000 species of freshwater and brackish gastropods. The total number of recent species of freshwater snails is about 4,000.
There are 444 Extinct species of gastropods (since the year 1500), 18 species Extinct in the Wild and 69 "Possibly Extinct" species.
The number of prehistoric (fossil) species of gastropods is at least 15,000 species.
Gastropods have a worldwide distribution from the near Arctic and Antarctic zones to the tropics. They have become adapted to almost every kind of existence on earth, having colonized every medium available except the air.
In habitats where there is not enough calcium carbonate to build a really solid shell, such as on some acidic soils on land, there are still various species of slugs, and also some snails with a thin translucent shell, mostly or entirely composed of the protein conchiolin.
Snails such as ''Sphincterochila boissieri'' and ''Xerocrassa seetzeni'' have adapted to desert conditions, other snails have adapted to an existence in ditches, near deepwater hydrothermal vents, the pounding surf of rocky shores, caves, and many other diverse areas.
thumb|The anatomy of an aquatic snail with a gill, a male prosobranch gastropod. Note that much of this anatomy does not apply to gastropods in other clades.Light yellow - bodyBrown - [[Gastropod shell|shell and operculumGreen - digestive systemLight purple - gillsYellow - osphradiumRed - heartPink - Dark violet - 1. foot 2. cerebral ganglion 3. pneumostome 4. upper commissura 5. osphradium 6. gills 7. pleural ganglion 8. atrium of heart 9. visceral ganglion 10. ventricle 11. foot 12. operculum 13. brain 14. mouth 15. tentacle (chemosensory, 2 or 4) 16. eye 17. penis (everted, normally internal) 18. esophageal nerve ring 19. pedal ganglion 20. lower commissura 21. vas deferens 22. pallial cavity / mantle cavity / respiratory cavity 23. parietal ganglion 24. anus 25. hepatopancreas 26. gonad 27. rectum 28. nephridium]]
Snails are distinguished by an anatomical process known as torsion, where the visceral mass of the animal rotates 180° to one side during development, such that the anus is situated more or less above the head. This process is unrelated to the coiling of the shell, which is a separate phenomenon. Torsion is present in all gastropods, but the opisthobranch gastropods are secondarily de-torted to various degrees.
Torsion occurs in two mechanistic stages. The first is muscular and the second is mutagenetic. The effects of torsion are primarily physiological - the organism develops an asymmetrical nature with the majority of growth occurring on the left side. This leads to the loss of right-paired appendages (e.g. ctenidia (comb-like respiratory apparatus), gonads, nephridia, etc.). Furthermore, the anus becomes redirected to the same space as the head. This is speculated to have some evolutionary function, as prior to torsion, when retracting into the shell, first the posterior end would get pulled in, and then the anterior. Now, the front can be retracted more easily, perhaps suggesting a defensive purpose.
However, this "rotation hypothesis" is being challenged by the "asymmetry hypothesis" in which the gastropod mantle cavity originated from one side only of a bilateral set of mantle cavities.
Gastropods typically have a well-defined head with two or four sensory tentacles with eyes, and a ventral foot, which gives them their name (Greek ''gaster'', stomach, and ''poda'', feet). The foremost division of the foot is called the propodium. Its function is to push away sediment as the snail crawls. The larval shell of a gastropod is called a protoconch.
Most shelled gastropods have a one piece shell, typically coiled or spiraled. This coiled shell usually opens on the right-hand side (as viewed with the shell apex pointing upward). Numerous species have an operculum, which in many species acts as a trapdoor to close the shell. This is usually made of a horn-like material, but in some molluscs it is calcareous. In the land slugs, the shell is reduced or absent, and the body is streamlined.
Lateral outgrowths on the body of nudibranchs are called cerata. These contain a part of digestive gland, which is called the diverticula.
Sensory organs of gastropods include olfactory organs, eyes, statocysts and mechanoreceptors. Gastropods have no hearing.
In terrestrial gastropods (land snails and slugs), the olfactory organs, located on the tips of the 4 tentacles, are the most important sensory organ, The chemosensory organs of opisthobranch marine gastropods are called rhinophores.
The majority of gastropods have simple visual organs, eye spots, that are situated either at the tip of the tentacles or the base of the tentacles. However "eyes" in gastropods range from these simple ocelli which cannot process an image being only able to distinguish light and dark, to more complex pit eyes, and even to lens eyes. In land snails and slugs, vision is not the most important sense, because they are mainly nocturnal animals.
The nervous system of gastropods includes the peripheral nervous system and the central nervous system. The central nervous system consist of ganglia connected by nerve cells. It includes paired ganglia: the cerebral ganglia, pedal ganglia, osphradial ganglia, pleural ganglia, parietal ganglia and the visceral ganglia. There are sometimes also buccal ganglia.
Many marine gastropods are burrowers, and have a siphon that extends out from the mantle edge. Sometimes the shell has a siphonal canal to accommodate this structure. A siphon enables the animal to draw water into their mantle cavity and over the gill. They use the siphon primarily to "taste" the water to detect prey from a distance. Gastropods with siphons tend to be either predators or scavengers.
In one large group of sea slugs, the gills are arranged as a rosette of feathery plumes on their backs, which gives rise to their other name, nudibranchs. Some nudibranchs have smooth or warty backs and have no visible gill mechanism, such that respiration may likely take place directly through the skin.
In many marine gastropods other than the opisthobranchs, there are separate sexes; most land gastropods however are hermaphrodites.
A few sea slugs are herbivores and some are carnivores. Many have distinct dietary preferences and regularly occur in close association with their food species.
Some predatory carnivorous gastropods include, for example: Cone shells, ''Testacella'', ''Daudebardia'', Ghost slug and others.
The first gastropods were exclusively marine, with the earliest representatives of the group appearing in the Late Cambrian (''Chippewaella'', ''Strepsodiscus''). Early Cambrian forms like ''Helcionella'' and ''Scenella'' are no longer considered gastropods, and the tiny coiled ''Aldanella'' of earliest Cambrian time is probably not even a mollusk. By the Ordovician period the gastropods were a varied group present in a range of aquatic habitats. Commonly, fossil gastropods from the rocks of the early Palaeozoic era are too poorly preserved for accurate identification. Still, the Silurian genus ''Poleumita'' contains fiftean identified species. Fossil gastropods were less common during the Palaeozoic era than bivalves.
Most of the gastropods of the Palaeozoic era belong to primitive groups, a few of which still survive today. By the Carboniferous period many of the shapes we see in living gastropods can be matched in the fossil record, but despite these similarities in appearance the majority of these older forms are not directly related to living forms. It was during the Mesozoic era that the ancestors of many of the living gastropods evolved.
One of the earliest known terrestrial (land-dwelling) gastropods is ''Maturipupa'', which is found in the Coal Measures of the Carboniferous period in Europe, but relatives of the modern land snails are rare before the Cretaceous period, when the familiar ''Helix'' first appeared.
In rocks of the Mesozoic era, gastropods are slightly more common as fossils, their shells are often well preserved. Their fossils occur in ancient beds deposited in both freshwater and marine environments. The "Purbeck Marble" of the Jurassic period and the "Sussex Marble" of the early Cretaceous period, which both occur in southern England, are limestones containing the tightly packed remains of the pond snail ''Viviparus''.
Rocks of the Cenozoic era yield very large numbers of gastropod fossils, many of these fossils being closely related to modern living forms. The diversity of the gastropods increased markedly at the beginning of this era, along with that of the bivalves.
Certain trail-like markings preserved in ancient sedimentary rocks are thought to have been made by gastropods crawling over the soft mud and sand. Although these trails are of debatable origin, some of them do resemble the trails made by living gastropods today.
Gastropod fossils may sometimes be confused with ammonites or other shelled cephalopods. An example of this is ''Bellerophon'' from the limestones of the Carboniferous period in Europe, the shell of which is planispirally coiled and can be mistaken for the shell of a cephalopod.
Gastropods are one of the groups that record the changes in fauna caused by the advance and retreat of the Ice Sheets during the Pleistocene epoch.
In the older classification of the gastropods, there were four subclasses:
The taxonomy of the Gastropoda is under constant revision, and more and more of the old taxonomy is being abandoned, as the results of DNA studies slowly become clearer. Nevertheless a few of the older terms such as "opisthobranch" and "prosobranch" are still sometimes used in a descriptive way.
New insights based on DNA sequencing of gastropods have produced some revolutionary new taxonomic insights. In the case of the Gastropoda, the taxonomy is now gradually being rewritten to embody strictly monophyletic groups (only one lineage of gastropods in each group). Integrating new findings into a working taxonomy will continue to be a challenge in coming years. Consistent ranks within the taxonomy at the level of subclass, superorder, order and suborder have already been abandoned as unworkable. Ongoing revisions of the higher taxonomic levels are to be expected in the near future.
Convergent evolution, which appears to exist at especially high frequency in the class Gastropoda class, may account for the observed differences between the older phylogenies which were based on morphological data, and more recent gene-sequencing studies.
Bouchet & Rocroi (2005) made sweeping changes in the systematics, resulting in a taxonomy that is a step closer to the evolutionary history of the phylum.
The Bouchet & Rocroi classification system is based partly on the older systems of classification, and partly on new cladistic research. In the past, the taxonomy of gastropods was largely based on phenetic morphological characters of the taxa. The recent advances are more based on molecular characters from DNA and RNA research. This has made the taxonomical ranks and their hierarchy controversial. The debate about these issues is not likely to end soon.
In the Bouchet, Rocroi ''et al.'' taxonomy, the authors have used unranked clades for taxa above the rank of superfamily (replacing the ranks suborder, order, superorder and subclass), while using the traditional Linnaean approach for all taxa below the rank of superfamily. Whenever monophyly has not been tested, or is known to be paraphyletic or polyphyletic, the term "group" or "informal group" has been used. The classification of families into subfamilies is often not well resolved, and should be regarded as the best possible hypothesis.
In 2004, Brian Simison and David R. Lindberg showed possible diphyletic origins of the Gastropoda based on mitochondrial gene order and amino acid sequence analyses of complete genes.
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This text is licensed under the Creative Commons CC-BY-SA License. This text was originally published on Wikipedia and was developed by the Wikipedia community.
Competing in only his second Formula Ford event, Ingall finished third in a support race at the 1988 Australian Grand Prix in Adelaide.
Over the next few years Ingall proved to be a force in the Australian Formula Ford Championship finishing runner-up before claiming the crown in the 1990.
After almost claiming the British Formula Ford Championship in 1991, Ingall headed to Europe in 1992 to drive for Opel Team Schiubel in the prestigious German Formula 3 Championship. Ingall also had the opportunity to compete at the Macau Grand Prix and surprised many people by starting 23rd and being in a position by mid-way through the race to overtake David Coulthard for fifth position.
Over the northern winter Ingall competed in the New Zealand Dunlop Formula Ford series, winning easily with 10 victories from 12 races.
Ingall made history in 1993 returning to Britain to drive for the factory Van Diemen team to win 13 out of the 16 races in the British Formula Ford Championship and in the process recording the highest number of wins in a single season in the history of Formula Ford.
The season was finished off by winning one of the most prestigious single-seater events - the Formula Ford Festival and World Cup at Brands Hatch in Britain.
Ingall was never able to live up to his full potential in Europe due to a lack of funding and later returned to Australia. This is a common tale for Australian race drivers in Europe.
During his first year, Ingall claimed his maiden victory at Calder Park and then went on to win the Bathurst 1000 for the second time in 1997.
In his seven years with Perkins Engineering, Ingall finish runner-up in the championship three times (1998, 1999 and 2001) and was third in 1997.
He also went on to win rounds at the Queensland 300 and the Gillette V8 Supercar Challenge at the Lexmark Indy 300 on the Gold Coast, Queensland before eventually finishing seventh outright in the championship.
In 2004 Ingall was again consistent with the highlight of the season coming with a round win at Symmons Plains in round 12. In the final round at Eastern Creek Raceway, he finished third overall to jump from fourth to second in the championship in his Caltex Ford Falcon and give Stone Brothers Racing an historic 1-2 finish as his teammate Marcos Ambrose won the championship.
In 2005 Ingall went one better, collecting his first V8 Supercar championship after having been runner-up four times. Ingall went into the season with a plan and he followed it to the final race of the season, which was held at the Phillip Island circuit. He raced “smarter” than he ever had before and worked out his strategy around the V8 Supercars points system to collect the title ahead of Craig Lowndes and Marcos Ambrose.
In 2006 Ingall saw the championship slip from his grasp due to poor performance from his car which saw him finish the championship in eighth place.
In the lead-up to the 2007 season finale, Ingall announced he was Holden bound, thus leaving Stone Brothers Racing and Ford after five years. He finished the championship for the first time outside the top 10, placing a disappointing eleventh. For 2008 Ingall will race at Paul Morris Motorsport and will be racing under the brand SuperCheap Auto Racing.
“If I didn’t stick it in (during) the warm up, the distributor drive would have probably gone in the warm-up so we would have found the problem before the race, so one thing led to another,” said Ingall. “At the end of the day it was all related to the shunt.”
In 2010, Ingall will be joined by New Zealander Greg Murphy.
Category:V8 Supercar drivers Category:British Formula Renault 2.0 drivers Category:German Formula Three Championship drivers Category:Japanese Formula Three Championship drivers Category:Australian racecar drivers Category:Australian amputees Category:1964 births Category:Living people Category:Bathurst 1000 winners Category:Formula Ford drivers Category:Australian Touring Car Championship drivers Category:Australian people of English descent
pl:Russell Ingall sv:Russell IngallThis text is licensed under the Creative Commons CC-BY-SA License. This text was originally published on Wikipedia and was developed by the Wikipedia community.
Greg Murphy (born 23 August 1972 in Hastings, New Zealand) is a racing driver, best known as a four-time winner of the Bathurst 1000. Greg Murphy joined Jeremy Clarkson and Richard Hammond presenting Top Gear, when it had its first international Live show at ASB Showgrounds in Auckland from February 12 - 15th 2009, and again when the show returned in 2010 with James May replacing Hammond.
In 1999, he joined Wynns Racing and paired with Steven Richards to win the Bathurst 1000. In the 2001 V8 Supercar season, Murphy joined the newly-formed Kmart Racing team and had two Bathurst 1000 wins with teammate Rick Kelly, in 2003 and 2004. Murphy had two championship runner-ups while at Kmart Racing, in 2002 and 2003. In 2001 and 2004 he finished fourth in the championship.
He is one of the best known V8 Supercar drivers (car No 51 since 2001) and has won four rounds at his home circuit at Pukekohe, near Auckland (2001, 2002, 2003 and 2005). At the Mount Panorama circuit at Bathurst, Murphy has the second best record for a Kiwi with four wins, compared to Jim Richards' seven.
Murphy's 2003 pole position of 2m.06.8594s at Bathurst stood as the fastest lap ever recorded at Mount Panorama until eclipsed seven years later,.
Murphy joined Tasman Motorsport in 2007.
Paul Morris Motorsport announced Murphy will be join the team in 2010 to drive Castrol supported Commodore. While the partnership looked like it was to reignite the "old murph" his performances and a poor car failed to produce results thus the announcement came in November 2010 that the partnership would split. Murphy joined Kelly Racing for 2011 in a very late deal.
Category:New Zealand racecar drivers Category:V8 Supercar drivers Category:1972 births Category:Living people Category:People from Hastings, New Zealand Category:Bathurst 1000 winners Category:24 Hours of Le Mans drivers Category:Australian Touring Car Championship drivers Category:Formula Holden drivers
sv:Greg MurphyThis text is licensed under the Creative Commons CC-BY-SA License. This text was originally published on Wikipedia and was developed by the Wikipedia community.
| name | Joaquín Rodrigo |
|---|---|
| birth date | November 22, 1901 |
| birth place | Sagunto, Valencia, Spain |
| death date | July 06, 1999 |
| death place | Madrid |
| nationality | Spanish |
| spouse | Victoria Kamhi |
| salary | }} |
Joaquín Rodrigo Vidre, 1st Marquis of the Gardens of Aranjuez (November 22, 1901July 6, 1999), commonly known as Joaquín Rodrigo, was a composer of classical music and a virtuoso pianist. Despite being nearly blind from an early age, he achieved great success. Rodrigo's music counts among some of the most popular of the 20th century, particularly his ''Concierto de Aranjuez'', considered one of the pinnacles of the Spanish music and guitar concerto repertoire.
Rodrigo studied music under Francisco Antich in Valencia and under Paul Dukas at the École Normale de Musique in Paris. After briefly returning to Spain, he went to Paris again to study musicology, first under Maurice Emmanuel and then under André Pirro. His first published compositions date from 1940. In 1943 he received Spain's National Prize for Orchestra for ''Cinco piezas infantiles'' ("Five Children's Pieces"), based on his earlier composition of the same piece for two pianos, premiered by Ricardo Viñes. From 1947 Rodrigo was a professor of music history, holding the Manuel de Falla Chair of Music in the Faculty of Philosophy and Letters, at Complutense University of Madrid.
His most famous work, ''Concierto de Aranjuez,'' was composed in 1939 in Paris, and in later life he and his wife declared that it was written as a response to the miscarriage of their first child. It is a concerto for guitar and orchestra. The central adagio movement is one of the most recognizable in 20th century classical music, featuring the interplay of guitar with English horn. This movement was later adapted by the conductor Gil Evans for Miles Davis' 1960 album Sketches of Spain. The Concerto was adapted by the composer himself for Harp and Orchestra and dedicated to Nicanor Zabaleta.
The success of this concerto led to commissions from a number of prominent soloists, including the flautist James Galway and the cellist Julian Lloyd Webber for whom Rodrigo composed his ''Concierto como un divertimento'' and ''Concierto serenata'' for Harp and Orchestra dedicated to Nicanor Zabaleta. In 1954 Rodrigo composed ''Fantasía para un gentilhombre'' at the request of Andrés Segovia. His ''Concierto Andaluz,'' for four guitars and orchestra, was commissioned by Celedonio Romero for himself and his three sons.
None of Rodrigo's works, however, achieved the popular and critical success of the ''Concierto de Aranjuez'' and the ''Fantasia para un gentilhombre''. These two works are very often paired in recordings.
He was awarded Spain's highest award for composition, the Premio Nacional de Música, in 1983. On 30 December 1991, Rodrigo was raised into the Spanish nobility by King Juan Carlos I with the hereditary title of ''Marqués de los Jardines de Aranjuez'' (English: Marquis of the Gardens of Aranjuez). He received the prestigious Prince of Asturias Award—Spain's highest civilian honor—in 1996. He was named Commander of the Order of Arts and Letters by the French government in 1998.
He married Victoria Kamhi, a Turkish-born pianist whom he had met in Paris, on 19 January 1933, in Valencia. Their daughter, Cecilia, was born 27 January 1941. Rodrigo died in 1999 in Madrid at the age of 97, and was succeeded as Marqués de los Jardines de Aranjuez by his daughter. Joaquín Rodrigo and his wife Victoria are buried at the cemetery at Aranjuez.
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Category:1901 births Category:1999 deaths Category:People from Valencia (province) Category:Blind musicians Category:Composers for the classical guitar Category:20th-century classical composers Category:Spanish composers Category:Commandeurs of the Ordre des Arts et des Lettres Category:Commanders Crosses of the Order of Merit of the Federal Republic of Germany
ar:خواكين رودريغو az:Xoakin Rodriqo bs:Joaquín Rodrigo bg:Хоакин Родриго ca:Joaquín Rodrigo Vidre cs:Joaquín Rodrigo cy:Joaquín Rodrigo da:Joaquín Rodrigo de:Joaquín Rodrigo et:Joaquín Rodrigo es:Joaquín Rodrigo eo:Joaquín Rodrigo fa:خواکین رودریگو fr:Joaquín Rodrigo hr:Joaquín Rodrigo io:Joaquín Rodrigo it:Joaquín Rodrigo he:חואקין רודריגו la:Ioaquim Rodrigo hu:Joaquín Rodrigo arz:خواكين رودريجو nl:Joaquín Rodrigo ja:ホアキン・ロドリーゴ no:Joaquín Rodrigo pl:Joaquín Rodrigo pt:Joaquín Rodrigo ru:Родриго, Хоакин simple:Joaquín Rodrigo sl:Joaquín Rodrigo sr:Хоакин Родриго fi:Joaquín Rodrigo sv:Joaquín Rodrigo tr:Joaquín Rodrigo uk:Хоакін Відре Родріго vi:Joaquín Rodrigo zh:霍亞金·羅德利果This text is licensed under the Creative Commons CC-BY-SA License. This text was originally published on Wikipedia and was developed by the Wikipedia community.
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